Saturday, 20 April 2024

Ardipithecus kadabba: The oldest member of the Human Family.

Fossil Hominid remains from the Late Miocene Middle Awash deposits. (a) ALAVP-2/10, mandible and all associated teeth; ALA-VP-2/120, ulna and humerus shaft; ALA-VP-2/11, hand phalanx . (b) AME-VP-1/71, lateral, plantar and dorsal views of foot phalanx. (c) STD-VP-2, teeth and partial clavicle. (d) DID-VP-1/80, hand phalanx. (e) ASKVP-3/160, occlusal, mesial and buccal views; ASK-VP-3/78, posterior view. All images are at the same scale. Haile-Selassie (2001).

In the late 1990s Ethiopian palaeoanthropologist Yohannes Haile-Selassie led a series of excavations at Alaya in the Middle Awash area of the Afar Region of Ethiopia. These uncovered an ancient woodland habitat, dated to between 5.2 and 5.8 million years ago, and yielded, among other things, a collection of loose teeth and bone fragments, which Haile-Selassie assigned to the Primate Ardipithecus, at that time believed to be an ancestor of the living Chimpanzees. At that time these Late Miocene fossils were the oldest known member of the Chimpanzee lineage, only slightly younger than the presumed split between the ancestors of Chimpanzees and Humans, between 6.5 and 5.5 million years ago. These specimens were older than and not identical to those previously assigned to the Pliocene Ardipithecus ramidus, causing Haile-Selassie to erect a new subspecies, Ardipithecus ramidus kadabba ('kadabba' implies the first ancestor in the Afar language).

Ethiopian palaeoanthropologist Yohannes Haile-Selassie in March 2017. Wikimedia Commons.

Further work on both the site and the fossils it yielded refined the age of the specimens to between 5.77 and 5.54 million years old. More notably, work on both Ardipithecus kadabba and Ardipithecus ramidus by Haile Salassie, along with American palaeoanthropologist Tim White, and Japanese palaeoanthropologist Gen Suwa, led to a revision of the genus Ardipithecus. Thus, Ardipithecus kadabba was promoted to a full species, probably ancestral to Ardipithecus ramidus (making it a 'chronospecies'). Furthermore, it is no longer considered to oldest ancestor of the Chimpanzee lineage, but instead is placed on the Hominin side of the family tree (anything more closely related to Humans than to Chimpanzees is a Hominin), making it the oldest member of the Human family.

Sunday, 14 April 2024

Java Man.

The original Trinil 1 (tooth), Trinil 2 (skullcap), and Trinil 3 (femur) specimens discovered by Eugene Dubois on the banks of the Bengawan Solo River in 1891-2, collectively refered to as 'Java Man'. Now in the collection of the Naturalis Biodiversity Center in Leidan, the Netherlands.. Peter Maas/Wikimedia Commons.

 In 1879, Charles Darwin's book 'The Descent of Man, and Selection in Relation to Sex' had proposed (amongst other things) that modern Humans were descended from African Apes. This was a matter of considerable controversy at the time, with many people, and in particular several religious groups, declaring the proposition highly offensive. Among the scientific community, and in particular those members of it who studied Human and animal anatomy, this was less of a surprise, and broadly in line with what many other people had been thinking. 

One aspect of the theory, which was widely contested, however, was the idea that Humanity had originated in Africa. In the nineteenth century, it was generally assumed that civilization had originated in Asia, and spread into Africa via Egypt, which contained the oldest known archaeological sites on the continent at that time. Since Asia is also home to Apes (including the Orangutan, which was then little understood, and many people believed could talk), it seemed quite possible that this continent could have been home to not just the origin of civilization, but the origin of Humanity itself.

Marie Eugène François Thomas Dubois, a trained physician with an interest in zoology working at the University of Amsterdam was inspired by this idea, and sought funding for an expedition to the Dutch East Indies (now Indonesia) to search for evidence of this origin. When this was (unsurprisingly) rejected, he enlisted as a surgeon with the Royal Netherlands East Indies Army, receiving a posting in Sumatra in 1887, where he began is search for a hypothetical Human ancestor.

Dubois's excavations in caves in Sumatra (inspired by the discovery of Neanderthal remains in caves at several locations in Europe), failed to produce any evidence of ancient Humans, but did uncover a range of large Mammal skeletons, bringing him to the attention of the colonial government. Palaeontology was still a new science in the late nineteenth century, and the discovery of large vertebrate fossils was a matter of national prestige. In Europe, an countries such as Britian, France, Germany, and even Belgium had all made impressive discoveries on their home territories, as had the young United States, and British and German colonies in Africa. Neither the Netherlands, nor any of its colonies, had produced any significant fossil discoveries at this time, making Dubois' discoveries significant for the colonial administration.

Marie Eugène François Thomas Dubois (Eugène Dubois). Wikimedia Commons.

In 1889 Dubois was relieved of his military duties, and provided with two assistants from the East Indies Army's corps of engineers, and fifty convict labourers. After another season in Sumatra, during which no Human remains were found, Dubois switched his attention to Java in 1890. This proved to be successful, with his team uncovering the molar and skullcap now known as Java Man in 1891, and the femur in 1892.

Dubois described the molar and skullcap under the name Anthropopithecus, a now obsolete term used to describe both some Hominin fossils (including a tooth from the Sivalik Hills of India, which resembled the one from Java), as well as the living Chimpanzee. He envisaged the living individual as being a Human-like Gibbon (his preferred choice for a Human ancestor) and living Humans. Following the discovery of the femur, which was longer and straighter than that of any Ape, emphasising the Human-like qualities of the remains, Dubois amended this to Anthropopithecus erectus, emphasising that the creature had an upright, Human-like stance. 

Dubois was originally cautious about his find, referring to it as a Human-like Ape, but refraining from designating it as a Human ancestor. However, examination of the skull-cap suggested that it had a cranial capacity of about 900 cm², far larger than any Ape, further emphasising the fossil's Human affinities, which led Dubois to redescribe the specimen as Pithecanthropus erectus, using a genus name the German naturalist had used for a (then hypothetical) missing link between Apes and Humans.

In the 1930s German palaeontologist Gustav Heinrich Ralph von Koenigswald carried out a series of further excavations, on Java, uncovering a number of additional Hominin specimens, which he believed to be closely related to those Dubois had discovered, and assigned them to the same species. Surprisingly, Dubois objected strongly to this, claiming that his fossils were those of Ape-men (and therefore a missing link between the two groups) while von Koenigswald's were close to modern Humans.

'Solo Man', one of the Hominin specimens discovered on Java by Gustav Heinrich Ralph von Koenigswald, now in the collection of the Smithsonian Natural History Museum. Ryan Somma/Wikimedia Commons.

Following the Second World War various scientists reviewed Dubois'  and von Koenigswald's material, as well as other Hominin fossils from other sites in Asia and Africa, coming to the conclusion that these all belonged to a single species, which German biologist Ernst Mayr named Homo erectus, placing it in the same genus as modern Humans, but using Dubois' specific name, which had priority as the first specimen described, thus making Java Man the holotype of the species. 

Today it is more-or-less universally accepted that Humanity has its roots in Africa, and Homo erectus is considered to have been the first Human species to have spread out of Africa, moving across Arabia and into South Asia, then China and Southeast Asia, eventually down into the Sundaland Region, which included Sumatra and Java, but which was still attached to mainland Asia during the Pleistocene, when sealevels were generally lower. 

Sundaland at the Last Glacial Maximum, showing the modern distribution of land in dark grey and the additional land exposed during the glacial maximum in light grey. The northern boundary of Sundaland defined by 9°N latitude shown as a dashed line. Possible lakes are marked by the letter L, the mouths of the major Molengraaff Rivers are indicated by letters as follows: 1, South Sunda River; 2, North Sunda River; 3, Siam River; 4, Malacca River. Bird et al. (2005).

The exact age range of Homo eructus is unclear; fossils dating from between about 1.4 million years ago and about 400 000 years ago have been assigned to the species, although there is considerable variation between these, leading some palaeoanthropologists to split the species into several different species or subspecies. As the holotype of Homo erectus, Java Man always remains within that species, sometimes being assigned to the subspecies Homo erectus erectus, while other specimens are re-assigned based upon their perceived similarity to it. This situation is not helped by the exact age of Java Man being unknown, not simply because dating methods were less advanced in the nineteenth century, but also because Dubois, having been provided with a free workforce by the authorities on Java, was not present when the fossils were excavated by his labourers, and was therefore uncertain about the horizon they were excavated from. 

However Homo erectus should be defined, the species is considered to have made remarkable advances over its predecessors, spreading out of Africa and across Asia into a range of entirely new environments, as well as developing the distinctive Acheulean tool set, distinguished by carefully shaped axes and points, unlike anything which had come before it. Homo erectus is also generally credited with having been the first Hominin to use fire, as well as having built the oldest wooden structures in Africa, half a million years ago.

Acheulean hand-axe from a hill-top plateau, 425 m above sealevel, 15 km to the northwest of the city of Naqada, Egypt. In the collection of the Petrie Museum in London. Osama Shukir Muhammed Amin/Wikimedia Commons.


Friday, 12 April 2024

Ruins of Kilwa Kisiwani and Ruins of Songo Mnara, UNESCO World Heritage Site, Tanzania.

The ruins of Kilwa Kisiwani, Tanzania. Ron Van Oers/UNESCO. 

Located on two islands close to each other just off the Tanzanian coast, about 300 km south of Dar es Salaam and 9° south of the Equator, are the remains of two port cites, Kilwa Kisiwani and Songo Mnara. The larger, Kilwa Kisiwani, was occupied from the 9th to the 19th century and reached its peak of prosperity in the 13th and 14th centuries. In 1331-1332, the great traveller, Ibn Battouta made a stop here and described Kilwa as one of the most beautiful cities of the world.

Kilwa Kisiwani and Songo Mnara were Swahili trading cities, and their prosperity was based on control of Indian Ocean trade with Arabia, India and China, particularly between the 13th and 16th centuries, when gold and ivory from the hinterland was traded for silver, carnelians, perfumes, Persian faience and Chinese porcelain. Kilwa Kisiwani minted its own currency in the 11th to 14th centuries. In the 16th century, the Portuguese established a fort on Kilwa Kisiwani and the decline of the two islands began.

Ruins of Kilwa Kisiwani, Tanzania. Wikimedia Commons.

The remains of Kilwa Kisiwani cover much of the island with many parts of the city still unexcavated. The substantial standing ruins, built of coral and lime mortar, include the Great Mosque constructed in the 11th century and considerably enlarged in the 13th century, and roofed entirely with domes and vaults, some decorated with embedded Chinese porcelain; the palace Husuni Kubwa built between about 1310 and 1333 with its large octagonal bathing pool; Husuni Ndogo, numerous mosques, the Gereza (prison) constructed on the ruins of the Portuguese fort and an entire urban complex with houses, public squares, burial grounds, etc.

The ruins of Songo Mnara, at the northern end of the island, consist of the remains of five mosques, a palace complex, and some thirty-three domestic dwellings constructed of coral stones and wood within enclosing walls.

Ruins of Songo Mnara, Tanzania. Ron Van Oers/UNESCO. 

The islands of Kilwa Kisiwani and Songo Mnara bear exceptional testimony to the expansion of Swahili coastal culture, the lslamization of East Africa and the extraordinarily extensive and prosperous Indian Ocean trade from the medieval period up to the modern era.